??>?? 8:????34567?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? ?R?fW?q_?T\???x?|?e?v?Q?|? ?xvzh?f\???|b_?v9e?S??YOb??|?v?N??X?R?~5%0 T?e ?\???v}|?|b_`_NN\???T(?o`o`?vsQ ?/f?T(??hKm-N?v???ch0?vMR(W\??'Y?R?gr?SO N???[MO0R?N?|b_?vsQ?vQTL?h?1-1 ?h?1-2 ??2015t^ ?Wu)R(uYanda1817/Beinong6M?u?vRIL?SO ?(W1B01D02A02B02D03B03D04A04B04D05A05B06B07A?S7B?gr?SO N?[MO0R32*N?|?QTL ?vQ-NS+T2A?gr?SO N?vN*N(WhQ萯s?X-N3z?[?v;NHe?|?QTL0(W5A06B07D?gr?SO N?[MO0R?|?[?GW ?QTL?Wu et al., 2015 ?0Mohler(uPamier/FormatM?u?vDH?SO ?(W1B02A02D04D?S5A?gr?SO N?[MO0R?|??v;NHeQTL0(W1B02A02B02D03A03B04D05B05D?T6A?gr?SO N?[MO0R10*N?|?[QTL0(W1B02A02D03A04A04D05A05B05D?gr?SO N?[MO0R?Nq_?T}|?|b??y?vQTL?Mohler et al., 2016 ?0Cheng)R(uWangshuibai/ Nanda2419M?u?vRIL?SO ?(W1A02A02D?T5A?gr?SO N?[MO0R?N4*N?|?QTL ??N(W5A?gr?SO N?hKm0R?NN*N?|?[QTL?Cheng et al., 2017 ?0UOwZwZI{?2017 ?)R(uhT??18?T?|q\?I?T015?N?]Tb?v?Nbc?|?SO ?(W2D?gr?SO N?hKm0R3z?[ T?eq_?TWz?|?0?|?0}|?|hT?I{'`?r?vYHe'`QTLMO?p0KumariO(uNW1014 /HUW468M?u?vRIL?SO ?(W1A06A02B?T 7B?gr?SO N?[MO0R?N5*N}|?|??[?k?GLWR ?QTL ?(W4A05A ?T7B?gr?SO N?[MO0R?N3*N}|?|hT??GPL ?QTL,(W5D07D?gr?SO N?[MO0R?N3*N?|b??y?GAS ?QTL ?(W4A05A?gr?SO N?[MO0R?N2*N?|?QTL ?(W6A?T7D?gr?SO N?[MO0R2*N?|?[QTL?Kumari et al., 2018 ?0Li)R(uDoumai ? Shi 4185?D ? S ?, Gaocheng 8901 ? Zhoumai 16?G ? Z ??TLinmai 2 ? Zhong 892?L ? Z ??g?^?vRIL?SO ??L??NhQ?W?V?~?N?'`?r??R?g ?(W1AL01BS01BL04BL04DS05BS05DL06AL06DL07AS?T7DS?gr?SO N?[MO0R11*N?|??vQTLMO?p0(W 2BL03AL03B04AS04AL04BS04DS05AL05BL05DL06AL07AL0?gr?SO N?[MO0R?N13*N?|?[QTLMO?p?Li et al., 2018 ?0 ,g???? ??S?Q'Y3753?SvQ?z?SSO564?g?^F2:3R?y?SO ??ǐ?g?^?RW? O?V1?v^?L?QTL?[MOR?g0?N?|b_'`?r:Nxvz?vh ??NR?[MO0?|?~?[MO0P ??W?VKQ???T?W?V?R??t??[I{B\b??L??-N?meQ0Wxvz0 h?1-1\???|?QTLsxvz?U\ ?|? ?SO{|?W h??{|?W ?gr?SO h??W?S_ Li et al?2015 ? RIL DArT/ SSR 1B,1D,4B,5B,6A 3.03-14.4% Sun et al?2008 ? RIL SSR 2A,5D,6A 8.6-26.4% Wu et al?2015 ? RIL SSR/SNP 1B,1D,2A,2B,2D,3B,3D,4A, 4B,4D,5A,5B,6B,7A,7B 2.62%-44.39% Mohler et al?2016 ? DH SSR/SNP 1B, 2A, 2D, 4D, 5A 1.0-23.8% Cheng et al?2017 ? RIL SSR 1A,2A,2D,5A 9.3-32.6% UOwZwZ?2017 ? BC1F SSR 2D,3D,7D 3.68-8.28% Li et al?2018 ? RIL SNP 1A, 1B, 4B, 4D, 5B, 5D, 6A, 6D, 7A ,7D 2.4-17.1% Kumari?2018 ? RIL SSR 4A,5A 53-53.84% Cui et al?2014 ? RIL SSR/ RAPD 1B,2A,2B,2D,3A,3B, 5A,6B,7A,7B 3.10-31.14% h?1-2\???|?[QTLsxvz?U\ ?|?[ ?SO{|?W h??{|?W ?gr?SO h??W?S_ Li et al?2015 ? RIL DArT/SSR 1B,3D,4B,5B,6A,6B 4.61-14.34% Sun et al?2008 ? RIL SSR 1A,1B,2B,4A,4B 6.3-12.1% Wu et al?2015 ? RIL SSR/SNP 5A,6B,7D 3.69% -12.30% Mohler?2016 ? DH SSR/SNP 1B,2A,2B,2D,3A, 3B, 4D, 5B, 5D,6A 1.2-17.9% Cheng et al?2017 ? RIL SSR 5A 13.6-19.2% UOwZwZI{?2017 ? BC1F7 SSR 3D 4.1766% Li et al?2018 ? RIL SNP 2B, 3A, 3B, 4A, 4B, 4D, 5A, 5B, 5D, 6A ,7A 2.8-25.2%  ?S??e.s? Cheng, R., Kong, Z., Zhang, L., et al. Mapping QTLs controlling kernel dimensions in a wheat inter-varietal RIL mapping population. Theor Appl,.0246>@h|????????????????????????q?c?W?cIIIcIc?h?P?h$e?>*CJ PJ\?h?P?h?i)CJ PJ\?h?P?h ?>*CJ PJ\?h?P?h$e?>*CJPJ\?aJh?P?h ?>*CJPJ\?aJh?P?h ?CJ PJ\?h?P?h?i)CJ,\?h?P?h ?CJ,\?h?P?h ?CJ4PJ\?h?P?h?i)CJ4\?h?P?h ?CJ4\?h?P?hDU?CJ4\?h?P?h ?CJ4PJ\?h?P?h ?\? jh?P?hDU?CJU\?aJ0246?? b  x ? ? ? ? ? ? ???????????????????$a$gd ?gd ??f??d??VDY]?f?^?gd ???d??VDY]?^?gd ? $?]?a$gd ?$ ?X ?]?a$gd ? $?r]?ra$gdDU?????   " 4 L ` b j p t v x ? ? ? ? ? ? ? ? ? ? ?     " 6 L v x ? ? ? ? ? ? ? ? ? ? ? ? ? ? ? ? ? ? ? ?????????????????????????????????????????????????????h?P?h$e?\?h?P?h ?CJ PJ\?aJ h?P?h ?\?h?P?h ?PJ\?h?P?h$e?0JCJ PJ\?#jh?P?h$e?>*CJ PJU\?h?P?h$e?>*CJ PJ\?h?P?h ?>*CJ PJ\?h?P?h ?CJ PJ\?8? ? ? ? ? ? ? ? ? ? ?  ????????j??vkd2$$If???0%X ?%3?0???????X ??????????????4?4? a? $$Ifa$ $$Ifa$ $ ?A?^?a$ ? ? ? ? ? ? ? ? ? ? ?        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"?@?????$??????0??????????????????????????????????????????????????????????????????????????Uh?P?hicS\?aJh?P?h ?KH\?aJh?P?h ?\?aJh?P?h$e?H*\?aJh?P?h$e?\?aJh?P?h$e?\?aJJ??  0$d?$G$Ifa$gd$e?K$?kd?$IfK$L$?l?  ?r? ????]??????? t??0? ?? ????U?????????????????????????4?4? la?p?2??????????ytuq? & 6 ???$d?$G$Ifa$gd$e?K$6 8 V ^ 0$d?$G$Ifa$gd$e?K$?kd?$IfK$L$?l?  ?r? ????]??????? t??0? ?? ????U?????????????????????????4?4? la?p?2??????????ytuq?^ f ? ? ? ????$d?$G$Ifa$gd$e?$d?$G$Ifa$gd$e?K$? ? ? 0*$If?kd}$IfK$L$?l?  ?r? ????]??????? t??0? ?? ????U?????????????????????????4?4? la?p?2??????????ytuq?? ? ? ? @??$???0?2?v???~???f?ΘИ????????????????$If $IfgdicSckde$$If???X X ?0???????X ??????????4?4? a? Genet, 2017,130(7):1405-1414. Cui, Fa, Zhao, Chunhua, Ding, Anming,I{. Construction of an integrative linkage map and QTL mapping of grain yield-related traits using three related wheat RIL populations. Theor Appl Genet, 2014, 127(3):659-675. Kumari, S., Jaiswal, V., Mishra, V.K. et al. QTL mapping for some grain traits in bread wheat (Triticum aestivum L.). Physiol Mol Biol Plants, 2018, 24:909. Li, F., Wen, W., He, Z. et al. Genome-wide linkage mapping of yield-related traits in three Chinese bread wheat populations using high-density SNP markers. Theor Appl Genet, 2018, 131(9):1903-1924. Li, Q., Zhang, Y., Liu, T., et al. Genetic analysis of kernel weight and kernel size in wheat (Triticum aestivum L.) using unconditional and conditional QTL mapping. Molecular Breeding, 2015. Lobell DB, Schlenker W, Costa-Roberts J. Climate trends and global crop production since 1980. Science, 2011, 333(6042):616-20. Mohler V, Albrecht T, Castell A., et al. Considering causal genes in the genetic dissection of kernel traits in common wheat. J Appl Genet, 2016, 57(4):467-476. Rosegrant M W, Agcaoili M. Global food demand, supply, and price prospects to 2010. International Food Policy Research Institute, Washington, D.C., USA. Simmonds J, Scott P, Brinton J, Mestre TC, et al. A splice acceptor site mutation in TaGW2-A1 increases thousand grain weight in tetraploid and hexaploid wheat through wider and longer grains. Theor Appl Genet, 2016, 129(6):1099-1112. Tilman D, Cassman KG, Matson PA, Naylor R, Polasky S. Agricultural sustainability and intensive production practices. Nature, 2002, 418(6898):671-677. Wu QH, Chen YX, Zhou SH, Fu L, Chen JJ, Xiao Y, et al. High-density genetic linkage map construction and QTL mapping of grain shape and size in the wheat population Yanda1817 ? Beinong6. PLoS One, 2015, 10(2):e0118144. UOwZwZ. \?? ??????D?~?gr?SOGr?k?Nbc?|?v?g?^?S}|?|'`?r?vQTL?[MO[D]. ?lWS'Yf[, 2017. ?N0?[?e??R?S?eHh(fnxT*N?s???[?W{Qf[uR?e???R?vwQSO`??S?8h?R?l) N ?0?[?e??R ,gxvz)R(uEMS?b/g??S?Q'Y3753 ??NvQT?N-Nt??[?Q3z?[W? O?v'Y?|?z?SSO 564 0?ǐ)R(u?z?SSONΑu?W?Q'Y3753Bg?N ??g?^F2:3R?y?SO ??N\???|b_:Nxvz'`?r?L?QTL?[MO ?T?~?L??c6R\???|b_?W?V?vKQ???T?R??xvz ?:N\?????y9eo??c?O?e`?0 1??W?N?Km?^penc_?SInDelh???TSTARPh?? ??[~b?S?NKN?Y`'` ??g?^W? 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